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We devised techniques to exploit this information: cross-calibration, in which node date calibrations are reused across a phylogeny, and cross-bracing, in which node date calibrations are formally linked in a hierarchical Bayesian model.We apply these methods to proteins with ancient duplications that have remained associated and originated from plastid and mitochondrial endosymbionts: the α and β subunits of ATP synthase and its relatives, and the elongation factor thermo unstable.Chloroplasts and mitochondria descended from bacterial ancestors, but the dating of these primary endosymbiosis events remains very uncertain, despite their importance for our understanding of the evolution of both bacteria and eukaryotes.All phylogenetic dating in the Proterozoic and before is difficult: Significant debates surround potential fossil calibration points based on the interpretation of the Precambrian microbial fossil record, and strict molecular clock methods cannot be expected to yield accurate dates over such vast timescales because of strong heterogeneity in rates.The methods yield reductions in dating uncertainty of 14–26% while only using date calibrations derived from phylogenetically unambiguous Phanerozoic fossils of multicellular plants and animals.Our results suggest that primary plastid endosymbiosis occurred ∼900 Mya and mitochondrial endosymbiosis occurred ∼1,200 Mya.Men and women are rotated to meet each other over a series of short "dates" usually lasting from three to eight minutes depending on the organization running the event.

After he answered my ad, we went on our first date, and it was fabulous.For example, the presence of recycled bricks at an archaeological site indicates the sequence in which the structures were built.Similarly, in geology, if distinctive granitic pebbles can be found in the sediment beside a similar granitic body, it can be inferred that the granite, after cooling, had been uplifted and eroded and therefore was not injected into the adjacent rock sequence.From top to bottom: Rounded tan domes of the Navajo Sandstone, layered red Kayenta Formation, cliff-forming, vertically jointed, red Wingate Sandstone, slope-forming, purplish Chinle Formation, layered, lighter-red Moenkopi Formation, and white, layered Cutler Formation sandstone.Photo from Glen Canyon National Recreation Area, Utah.Although with clever detective work many complex time sequences or relative ages can be deduced, the ability to show that objects at two separated sites were formed at the same time requires additional information.